C. Young, M. Sewell, and M. Rice, Atlas of Marine Invertebrate Larvae, Aquaculture Research, vol.34, issue.5, 2002.
DOI : 10.1046/j.1365-2109.2003.00819.x

T. Turk and W. Kem, The phylum Cnidaria and investigations of its toxins and venoms until 1990, Toxicon, vol.54, issue.8, pp.1031-1037, 2009.
DOI : 10.1016/j.toxicon.2009.06.031

A. Collins, P. Schuchert, A. Marques, T. Jankowski, M. Medina et al., Medusozoan Phylogeny and Character Evolution Clarified by New Large and Small Subunit rDNA Data and an Assessment of the Utility of Phylogenetic Mixture Models, Systematic Biology, vol.55, issue.1, pp.97-115, 2006.
DOI : 10.1080/10635150500433615

L. Miranda, A. Collins, and A. Marques, Molecules Clarify a Cnidarian Life Cycle ??? The ???Hydrozoan??? Microhydrula limopsicola Is an Early Life Stage of the Staurozoan Haliclystus antarcticus, PLoS ONE, vol.5, issue.4, p.20418959, 2010.
DOI : 10.1371/journal.pone.0010182.t003

A. Marques, V. Haddad, E. Jr, and A. Migotto, Envenomation by a benthic Hydrozoa (Cnidaria): the case of Nemalecium lighti (Haleciidae), Toxicon, vol.40, issue.2, pp.213-215, 2002.
DOI : 10.1016/S0041-0101(01)00196-9

A. Collins, Phylogeny of Medusozoa and the evolution of cnidarian life cycles, Journal of Evolutionary Biology, vol.232, issue.217, pp.418-432, 2002.
DOI : 10.1046/j.1420-9101.2002.00403.x

B. Aerne, H. Gröger, P. Schuchert, J. Spring, and V. Schmid, The polyp and its medusa: a molecular approach, Sci Mar, vol.60, pp.7-16, 1996.

M. Daly, M. Brugler, P. Cartwright, A. Collins, M. Dawson et al., The phylum Cnidaria: A review of phylogenetic patterns and diversity 300 years after Linnaeus, Zootaxa, vol.1668, pp.127-182, 2007.

C. Lucas, W. Graham, C. Widmer, and L. Michael, Chapter Three?Jellyfish life histories: Role of polyps in forming and maintaining Scyphomedusa populations Advances in Marine Biology, pp.133-196, 2012.

B. Galliot and V. Schmid, Cnidarians as a model system for understanding evolution and regeneration, Int. J. Dev. Biol, vol.46, pp.39-48, 2002.

U. Technau and R. Steele, Evolutionary crossroads in developmental biology: Cnidaria Development 138, pp.1447-1458, 2011.
DOI : 10.1242/dev.048959
URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3062418

P. Cartwright and A. Nawrocki, Character Evolution in Hydrozoa (phylum Cnidaria), Integrative and Comparative Biology, vol.50, issue.3, pp.456-472, 2010.
DOI : 10.1093/icb/icq089

N. Gravier-bonnet and M. Lebon, (Cnidaria, Hydrozoa, Sertulariidae) from R??union Island, Indian Ocean, Invertebrate Reproduction & Development, vol.65, issue.1-3, pp.101-108, 2002.
DOI : 10.1080/00222939800770011

F. Boero and N. Gravier-bonnet, The life cycle of Pteroclava krempfi (Cnidaria, Hydrozoa, Cladocorynidae ), with notes on Asyncoryne philippina (Asyncorynidae), Sci Mar, vol.56, pp.65-76, 1995.

J. Bouillon and F. Boero, Phylogeny and classification of Hydroidomedusae. Amaltea edizioni, issue 24 of Thalassia Salentina, 2000.

L. Suescún-bolívar, R. Iglesias-prieto, and P. Thome, Induction of Glycerol Synthesis and Release in Cultured Symbiodinium, PLoS ONE, vol.7, issue.10, p.23071753, 2012.
DOI : 10.1371/journal.pone.0047182.t003

P. Ganot, A. Moya, V. Magnone, D. Allemand, P. Furla et al., Adaptations to Endosymbiosis in a Cnidarian-Dinoflagellate Association: Differential Gene Expression and Specific Gene Duplications, PLoS Genetics, vol.52, issue.7, p.21811417, 2011.
DOI : 10.1371/journal.pgen.1002187.s011

S. Davy, D. Allemand, and V. Weis, Cell Biology of Cnidarian-Dinoflagellate Symbiosis, Microbiology and Molecular Biology Reviews, vol.76, issue.2, pp.229-261, 2012.
DOI : 10.1128/MMBR.05014-11

E. Meyer and V. Weis, Study of Cnidarian-Algal Symbiosis in the ???Omics??? Age, The Biological Bulletin, vol.223, issue.1, pp.44-65, 2012.
DOI : 10.1086/BBLv223n1p44

M. Goodson, L. Whitehead, and A. Douglas, Symbiotic dinoflagellates in marine Cnidaria: diversity and function, Hydrobiologia, vol.461, issue.1/3, pp.79-82, 2001.
DOI : 10.1023/A:1012766609448

C. Ferrier-pages, J. Gattuso, G. Cauwet, J. Jaubert, and D. Allemand, Release of dissolved organic carbon and nitrogen by the zooxanthellate coral Galaxea fascicularis, Marine Ecology Progress Series, vol.172, pp.265-274, 1998.
DOI : 10.3354/meps172265

A. Venn, J. Loram, and A. Douglas, Photosynthetic symbioses in animals, Journal of Experimental Botany, vol.59, issue.5, pp.1069-1080, 2008.
DOI : 10.1093/jxb/erm328

C. Krediet, K. Ritchie, V. Paul, and M. Teplitski, Coral-associated micro-organisms and their roles in promoting coral health and thwarting diseases, Proceedings of the Royal Society B: Biological Sciences, vol.23, issue.7, p.20122328, 2013.
DOI : 10.1016/j.tree.2008.03.004

M. Burriesci, T. Raab, and J. Pringle, Evidence that glucose is the major transferred metabolite in dinoflagellate-cnidarian symbiosis, Journal of Experimental Biology, vol.215, issue.19, pp.3467-3477, 2012.
DOI : 10.1242/jeb.070946

D. Fautin, Structural diversity, systematics, and evolution of cnidae, Toxicon, vol.54, issue.8, pp.1054-1064, 2009.
DOI : 10.1016/j.toxicon.2009.02.024

J. Stachowicz and N. Lindquist, Hydroid defenses against predators: the importance of secondary metabolites versus nematocysts, Oecologia, vol.124, issue.2, pp.280-288, 2000.
DOI : 10.1007/s004420000372

T. Bosch, What Hydra Has to Say About the Role and Origin of Symbiotic Interactions, The Biological Bulletin, vol.223, issue.1, pp.78-84, 2012.
DOI : 10.1086/BBLv223n1p78

H. Kawaida, K. Ohba, Y. Koutake, H. Shimizu, H. Tachida et al., Symbiosis between hydra and chlorella: Molecular phylogenetic analysis and experimental study provide insight into its origin and evolution, Molecular Phylogenetics and Evolution, vol.66, issue.3, pp.906-914, 2013.
DOI : 10.1016/j.ympev.2012.11.018

F. Wilkerson, Bacterial symbionts on green hydra and their effect on phosphate uptake, Microbial Ecology, vol.262, issue.1, pp.85-92, 1980.
DOI : 10.1007/BF02020377

E. Verde and L. Mccloskey, Production, respiration, and photophysiology of the mangrove jellyfish Cassiopea xamachana symbiotic with zooxanthellae:effect of jellyfish size and season, Marine Ecology Progress Series, vol.168, pp.147-162, 1998.
DOI : 10.3354/meps168147

H. Felbeck, J. Childress, and G. Somero, Calvin-Benson cycle and sulphide oxidation enzymes in animals from sulphide-rich habitats, Nature, vol.281, issue.5830, pp.291-293, 1981.
DOI : 10.1038/293291a0

C. Cavanaugh, Z. Mckiness, I. Newton, and F. Stewart, Marine chemosynthetic symbioses. The prokaryotes, pp.475-507, 2006.

F. Stewart, I. Newton, and C. Cavanaugh, Chemosynthetic endosymbioses: adaptations to oxic???anoxic interfaces, Trends in Microbiology, vol.13, issue.9, pp.439-448, 2005.
DOI : 10.1016/j.tim.2005.07.007

N. Dubilier, C. Bergin, and C. Lott, Symbiotic diversity in marine animals: the art of harnessing chemosynthesis, Nature Reviews Microbiology, vol.214, issue.10, pp.725-740, 2008.
DOI : 10.1264/jsme2.22.136

M. Yamamoto and K. Takai, Sulfur Metabolisms in Epsilon- and Gamma-Proteobacteria in Deep-Sea Hydrothermal Fields, Frontiers in Microbiology, vol.2, p.21960986, 0192.
DOI : 10.3389/fmicb.2011.00192

K. Vopel, D. Thistle, J. Ott, M. Bright, and H. Roy, S flux sustains a chemoautotrophic symbiosis, Limnology and Oceanography, vol.50, issue.1, pp.128-133, 2005.
DOI : 10.4319/lo.2005.50.1.0128

T. Fenchel and R. Riedl, The sulfide system: a new biotic community underneath the oxidized layer of marine sand bottoms, Marine Biology, vol.7, issue.3, pp.255-268, 1970.
DOI : 10.4319/lo.1961.6.3.0262

T. Woyke, H. Teeling, N. Ivanova, M. Huntemann, M. Richter et al., Symbiosis insights through metagenomic analysis of a microbial consortium, Nature, vol.5, issue.7114, pp.950-955, 2006.
DOI : 10.1038/nature05192

V. Edgcomb, S. Breglia, N. Yubuki, D. Beaudoin, D. Patterson et al., Identity of epibiotic bacteria on symbiontid euglenozoans in O2-depleted marine sediments: evidence for symbiont and host co-evolution, The ISME Journal, vol.9, issue.2, pp.231-243, 2011.
DOI : 10.1038/ismej.2010.121

U. Hentschel, E. Berger, M. Bright, H. Felbeck, and J. Ott, Metabolism of nitrogen and sulfur in ectosymbiotic bacteria of marine nematodes (Nematoda, Stilbonematinae), Marine Ecology Progress Series, vol.183, pp.149-158, 1999.
DOI : 10.3354/meps183149

G. Tokuda, A. Yamada, K. Nakano, N. Arita, and H. Yamasaki, Colonization of Sulfurovum sp. on the gill surfaces of Alvinocaris longirostris, a deep-sea hydrothermal vent shrimp, Marine Ecology, vol.17, issue.1, pp.106-114, 2008.
DOI : 10.3354/meps284237

J. Bauermeister, K. Assig, and S. Dattagupta, Exploring the sulfide tolerance of ectosymbiotic Niphargus amphipods from the Frasassi caves, central Italy, Int J Speleol, vol.42, p.6, 2013.

J. Deming, A. Reysenbach, S. Macko, and C. Smith, Evidence for the microbial basis of a chemoautotrophic invertebrate community at a whale fall on the deep seafloor: Bone-colonizing bacteria and invertebrate endosymbionts, Microscopy Research and Technique, vol.131, issue.2, pp.162-170, 1997.
DOI : 10.1002/(SICI)1097-0029(19970415)37:2<162::AID-JEMT4>3.0.CO;2-Q

A. Girnth, S. Grünke, A. Lichtschlag, J. Felden, K. Knittel et al., A novel, mat-forming Thiomargarita population associated with a sulfidic fluid flow from a deep-sea mud volcano, Environmental Microbiology, vol.14, issue.2, pp.495-505, 2011.
DOI : 10.1111/j.1462-2920.2010.02353.x

R. Schauer, H. Røy, N. Augustin, H. Gennerich, M. Peters et al., Bacterial sulfur cycling shapes microbial communities in surface sediments of an ultramafic hydrothermal vent field, Environmental Microbiology, vol.62, issue.10, p.21895907, 2011.
DOI : 10.1111/j.1462-2920.2011.02530.x

M. Jean, S. Gonzalez-rizzo, P. Gauffre-autelin, S. Lengger, S. Schouten et al., Description of two new Beggiatoa species inhabiting marine mangrove sediments from Caribbean, 2015.

C. Fisher, Chemoautotrophic and methanotrophic symbioses in marine-invertebrates, Rev Aquat Sci, vol.2, pp.399-436, 1990.

V. Tarasov, A. Gebruk, A. Mironov, and L. Moskalev, Deep-sea and shallow-water hydrothermal vent communities: Two different phenomena?, Chemical Geology, vol.224, issue.1-3, pp.5-39, 2005.
DOI : 10.1016/j.chemgeo.2005.07.021

C. Berg and P. Alatalo, Potential of chemosynthesis in molluscan mariculture, Aquaculture, vol.39, issue.1-4, pp.165-179, 1984.
DOI : 10.1016/0044-8486(84)90264-3

M. Laurent, O. Gros, J. Brulport, F. Gaill, and N. Bris, Sunken wood habitat for thiotrophic symbiosis in mangrove swamps, Marine Environmental Research, vol.67, issue.2, pp.83-88, 2009.
DOI : 10.1016/j.marenvres.2008.11.006
URL : https://hal.archives-ouvertes.fr/hal-00751649

M. Laurent, L. Bris, N. Gaill, F. Gros, and O. , Dynamics of wood fall colonization in relation to sulfide concentration in a mangrove swamp, Marine Environmental Research, vol.87, issue.88, pp.85-95, 2013.
DOI : 10.1016/j.marenvres.2013.03.007

L. Frenkiel, O. Gros, and M. Mouëza, Gill structure in Lucina pectinata (Bivalvia: Lucinidae) with reference to hemoglobin in bivalves with symbiotic sulphur-oxidizing bacteria, Mar Biol, vol.125, pp.511-524, 1996.

M. Bauer-nebelsick, C. Bardele, and J. Ott, Redescription of Zoothamnium niveum (Hemprich & Ehrenberg, 1831) Ehrenberg, 1838 (Oligohymenophora, Peritrichida), a ciliate with ectosymbiotic, chemoautotrophic bacteria, European Journal of Protistology, vol.32, issue.1, pp.18-30, 1996.
DOI : 10.1016/S0932-4739(96)80036-8

D. Himmel, L. Maurin, O. Gros, and J. Mansot, Raman microspectrometry sulfur detection and characterization in the marine ectosymbiotic nematode Eubostrichus dianae (Desmodoridae, Stilbonematidae), Biology of the Cell, vol.284, issue.(3), pp.43-54, 2009.
DOI : 10.1042/BC20080051
URL : https://hal.archives-ouvertes.fr/hal-00753017

L. Maurin, D. Himmel, J. Mansot, and O. Gros, Raman microspectrometry as a powerful tool for a quick screening of thiotrophy: An application on mangrove swamp meiofauna of Guadeloupe (F.W.I.), Marine Environmental Research, vol.69, issue.5, 2010.
DOI : 10.1016/j.marenvres.2010.02.001
URL : https://hal.archives-ouvertes.fr/hal-00715329

D. Lane, B. Pace, G. Olsen, D. Stahl, M. Sogin et al., Rapid determination of 16S ribosomal RNA sequences for phylogenetic analyses., Proceedings of the National Academy of Sciences, vol.82, issue.20, pp.6955-6959, 1985.
DOI : 10.1073/pnas.82.20.6955

G. Giribet, S. Carranza, J. Baguñà, M. Riutort, and C. Ribera, First molecular evidence for the existence of a Tardigrada + Arthropoda clade, Molecular Biology and Evolution, vol.13, issue.1, pp.76-84, 1996.
DOI : 10.1093/oxfordjournals.molbev.a025573

S. Altschul, W. Gish, W. Miller, E. Myers, and D. Lipman, Basic local alignment search tool, Journal of Molecular Biology, vol.215, issue.3, pp.403-410, 1990.
DOI : 10.1016/S0022-2836(05)80360-2

K. Tamura, D. Peterson, N. Peterson, G. Stecher, M. Nei et al., MEGA5: Molecular Evolutionary Genetics Analysis Using Maximum Likelihood, Evolutionary Distance, and Maximum Parsimony Methods, Molecular Biology and Evolution, vol.28, issue.10, pp.2731-2739, 2011.
DOI : 10.1093/molbev/msr121

M. Novo, C. Gayoso, F. Pomar, M. Lucas, R. Barcelo et al., Sulphur accumulation after Verticillium dahliae infection of two pepper cultivars differing in degree of resistance, Plant Pathology, vol.63, issue.6, pp.998-1004, 2007.
DOI : 10.1104/pp.128.1.150

F. Muller, T. Brissac, L. Bris, N. Felbeck, H. Gros et al., First description of giant Archaea (Thaumarchaeota) associated with putative bacterial ectosymbionts in a sulfidic marine habitat, Environmental Microbiology, vol.43, issue.8, pp.2371-2383, 2010.
DOI : 10.1111/j.1462-2920.2010.02309.x

C. Rinke, S. Schmitz-esser, K. Stoecker, A. Nussbaumer, D. Molnár et al., "Candidatus Thiobios zoothamnicoli," an Ectosymbiotic Bacterium Covering the Giant Marine Ciliate Zoothamnium niveum, Applied and Environmental Microbiology, vol.72, issue.3, pp.2014-2021, 2006.
DOI : 10.1128/AEM.72.3.2014-2021.2006

S. Fraune and T. Bosch, Long-term maintenance of species-specific bacterial microbiota in the basal metazoan Hydra, Proceedings of the National Academy of Sciences, vol.104, issue.32, pp.13146-13151, 2007.
DOI : 10.1073/pnas.0703375104

C. Schuett and H. Doepke, Endobiotic bacteria and their pathogenic potential in cnidarian tentacles, Helgoland Marine Research, vol.44, issue.6776, pp.205-212, 2012.
DOI : 10.1007/s10152-009-0179-2
URL : https://hal.archives-ouvertes.fr/hal-00541672

D. Camillo, C. Luna, G. Bo, M. Giordano, G. Corinaldesi et al., Biodiversity of Prokaryotic Communities Associated with the Ectoderm of Ectopleura crocea (Cnidaria, Hydrozoa), PLoS ONE, vol.124, issue.6, p.22768172, 2012.
DOI : 10.1371/journal.pone.0039926.s001

R. Forest, S. Victor, A. Farooq, and K. Nancy, Diversity and distribution of coral-associated bacteria, Mar Ecol Prog Ser, vol.243, pp.1-10, 2002.

L. Reshef, O. Koren, Y. Loya, I. Zilber-rosenberg, and E. Rosenberg, The Coral Probiotic Hypothesis, Environmental Microbiology, vol.273, issue.12, pp.2068-2073, 2006.
DOI : 10.2307/1313569

O. Koren and E. Rosenberg, Bacteria Associated with Mucus and Tissues of the Coral Oculina patagonica in Summer and Winter, Applied and Environmental Microbiology, vol.72, issue.8, pp.5254-16885273, 2006.
DOI : 10.1128/AEM.00554-06

H. Ducklow and R. Mitchell, Bacterial populations and adaptations in the mucus layers on living corals1, Limnology and Oceanography, vol.24, issue.4, pp.715-725, 1979.
DOI : 10.4319/lo.1979.24.4.0715

C. Carlos, T. Torres, and L. Ottoboni, Bacterial communities and species-specific associations with the mucus of Brazilian coral species, Scientific Reports, vol.4, 2013.
DOI : 10.1038/srep01624

J. Costello, Laboratory culture and feeding of the hydromedusa Cladonema californicum Hyman (Anthomedusa: Cladonemidae), Journal of Experimental Marine Biology and Ecology, vol.123, issue.2, pp.177-188, 1988.
DOI : 10.1016/0022-0981(88)90168-2

J. Costello, Complete carbon and nitrogen budgets for the hydromedusaCladonema californicum (Anthomedusa: Cladonemidae), Marine Biology, vol.45, issue.1, pp.119-128, 1991.
DOI : 10.1007/BF01313479

J. Costello, Physiological response of the hydromedusa Cladonema californicum Hyman (Anthomedusa: Cladonemidae) to starvation and renewed feeding, Journal of Experimental Marine Biology and Ecology, vol.225, issue.1, pp.13-28, 1998.
DOI : 10.1016/S0022-0981(97)00204-9

J. Rees, The hydrozoan Cladonema in California: a possible introduction from East Asia, Pac Sci, vol.36, pp.439-444, 1982.

J. Grzymski, A. Murray, B. Campbell, M. Kaplarevic, G. Gao et al., Metagenome analysis of an extreme microbial symbiosis reveals eurythermal adaptation and metabolic flexibility, Proceedings of the National Academy of Sciences, vol.105, issue.45, pp.17516-17521, 2008.
DOI : 10.1073/pnas.0802782105

A. Bates, T. Harmer, G. Roeselers, and C. Cavanaugh, Phylogenetic Characterization of Episymbiotic Bacteria Hosted by a Hydrothermal Vent Limpet (Lepetodrilidae, Vetigastropoda), The Biological Bulletin, vol.220, issue.2, pp.118-127, 2011.
DOI : 10.1086/BBLv220n2p118

A. Moya, J. Peretó, R. Gil, and A. Latorre, Learning how to live together: genomic insights into prokaryote???animal symbioses, Nature Reviews Genetics, vol.34, issue.3, pp.218-229, 2008.
DOI : 10.1038/nrg2319

J. Chaston and H. Goodrich-blair, Common trends in mutualism revealed by model associations between invertebrates and bacteria: Table 1, FEMS Microbiology Reviews, vol.34, issue.1, pp.41-58, 2010.
DOI : 10.1111/j.1574-6976.2009.00193.x

J. Kimbell and M. Mcfall-ngai, The Squid-Vibrio Symbioses: From Demes to Genes, Integrative and Comparative Biology, vol.43, issue.2, pp.254-260, 2003.
DOI : 10.1093/icb/43.2.254

M. Mcfall-ngai, Unseen Forces: The Influence of Bacteria on Animal Development, Developmental Biology, vol.242, issue.1, pp.1-14, 2002.
DOI : 10.1006/dbio.2001.0522

C. Ruehland and N. Dubilier, Gamma- and epsilonproteobacterial ectosymbionts of a shallow-water marine worm are related to deep-sea hydrothermal vent ectosymbionts, Environmental Microbiology, vol.46, pp.2312-2326, 2010.
DOI : 10.1111/j.1462-2920.2010.02256.x

S. Dattagupta, I. Schaperdoth, A. Montanari, S. Mariani, N. Kita et al., A novel symbiosis between chemoautotrophic bacteria and a freshwater cave amphipod, The ISME Journal, vol.1, issue.8, pp.935-943, 2009.
DOI : 10.3354/meps187059

J. Ponsard, M. Cambon-bonavita, M. Zbinden, G. Lepoint, A. Joassin et al., Inorganic carbon fixation by chemosynthetic ectosymbionts and nutritional transfers to the hydrothermal vent host-shrimp Rimicaris exoculata, The ISME Journal, vol.121, issue.1, pp.96-109, 2013.
DOI : 10.1016/j.jembe.2008.03.009
URL : https://hal.archives-ouvertes.fr/hal-00808528